The Wolves of Isle Royale

Michigan Tech Press Releases 2000-2004

Go back to Ben Kilpela's Isle Royale Home Page


For the official web site of the Moose-Wolf Study, go to: http://www.isleroyalewolf.org/

Also, for the complete annual Moose-Wolf reports from 1996-2004, go to: http://www.dartmouth.edu/~chance/teaching_aids/books_articles/WolfReport.html


This Year's Moose-Wolf Study Report: 2004

Moose Count: just 750 in 2004, down from 900 in 2003.

Wolf Count: 29 in 2004, up from 18 in 2003.

The Story:

Wolves are up and moose are down this spring at Isle Royale National Park, the home of a 46-year study of predators and their prey.  Researchers suspect that a global warming trend may be behind the shift.

The moose population has slid to 750 on this Lake Superior wilderness island park, down from 900 last year and 1,100 in 2002. In the meantime, the number of wolves has seesawed upward over the past decade and is now up to 29, as many as the park has seen since 1980 and 11 more than last year.

What's bad for moose has been good for the wolves, and moose throughout North America have been hit hard by warmer temperatures that began in 1998 with El Nino and never let up, according to Professor Rolf Peterson of Michigan Technological University, who has lead the study of Isle Royale's wolves and moose for 34 years.

"What we think is happening is that wolves are cashing in on moose vulnerability that's been induced by a warmer climate," Peterson said.  The moose population has been stressed by higher temperatures, particularly the drought of 1998 and then warm fall of 2001. "Moose can't feed in the summertime if it's too hot," Peterson said. "They have a big fur coat on, and they can't sweat. They just sit in the shade or in the water."

When moose don't eat enough in summer, they can become weak, sickly and easy prey for wolves during the winter.  And heat precipitates another blight for the big herbivores: ticks.  "Warm weather in spring and fall leads to ticks the following winter, and ticks can kill moose," Peterson said. A single moose can be host to tens of thousands at a time, several per square inch, and each tick can suck up about a cubic centimeter of blood. Rather than browse, the moose scratch themselves against trees or bite their hair out trying to remove the parasites. Weight and blood loss often prove such a handicap that the moose don't survive.

As the moose population struggles against the heat and ticks, the wolves have thrived, largely because it's been easier for them to bring down their biggest prey.  "The wolves are killing about twice as many moose as they did last year," Peterson says, which allows them to maintain their peak population.

Initially, researchers didn't know what warmer temperatures would mean for Isle Royale's wolves and moose.  "In this region the change has involved warmer winters, especially in the late 1990s and early years of this century," Peterson said. "We couldn't anticipate the effect for moose, because warmer winters mean less snow and more tree growth, which helps them.  But it also leads to more ticks, and it impacts their feeding. With two pluses and two minuses, there was no way to forecast how it would come out in the wash. But it looks like it might be to the detriment of moose."

As to what has been causing the warmer temperatures, Peterson cites climate models presented in the journal Science that suggest that greenhouse gases may be the culprit. While researchers do not have proof that increasing levels of carbon dioxide and other gases are warming the Earth's atmosphere, the best computer models of climate can only account for such higher temperatures if greenhouse gases are included.

Peterson's study is funded by the National Park Service, the National Science Foundation and Earthwatch. 

Contact information: Rolf Peterson, 906-487-2179, ropeters@mtu.edu; Marcia Goodrich, Michigan Tech media relations manager, 906-487-2343, mtunews@mtu.edu

3/08/04, From Michigan Tech News


Last Year's Report: 2003

Moose Count: 900 this year, down from 1100 in 2002.

Wolf Count: 19 this year, up from 17 in 2002.

The Story:

Ticks Reduce Isle Royale Moose Population

A tick infestation accounts for the changes in the annual wolf-moose survey at Isle Royale National Park, according to Michigan Technological University's Rolf Peterson.

Peterson, a professor of wildlife ecology at Michigan Tech, said the 2003 survey counted 19 wolves on the island, as opposed to 17 last year. The island's moose population decreased from about 1,100 last year to 900 in 2003.

"Winter ticks will slow the moose down for a couple of years,” Peterson said. "The tick infestation resulted from an early, warm spring in 2001 and an unusually mild autumn that year. This was an event that affected moose herds from New Hampshire to Alberta since 2002. It has killed a lot of moose in the region.”

The Isle Royale wolf-moose survey is the longest running predator-prey study in the world, now in its 45th year. Peterson has conducted the study for the last 33 years. As an island in the middle of Lake Superior, Isle Royale presents a unique opportunity for such research.

“Our objective is to understand how the moose and wolves interact when humans are not affecting the outcome,” he said.  Peterson said there is a lag time before the moose feel the effects of the tick infestation. The ticks attach themselves to the moose in the fall and are on the moose all winter.

“The biggest effects are in late winter as female ticks withdraw blood to support their own reproduction in the spring, when they drop off the moose,” Peterson said.  Weather continues to play a role in another way. Isle Royale “had as long a stretch of cold as I’ve ever seen,” Peterson said. “Plus, there was no snow at all when we got there in early January.” In the two months he spent on the island, he reports a snow depth of no more than 10 inches.

The lack of snow benefits the moose, because they can freely roam the island. A larger snowfall causes the moose to hunker down in conifer swamps, making them easier prey for the wolves. Typically the wolves attack old moose and young calves.

"Many moose died in 1996,” he said, “so the average moose is only five or six years old. There a lot of young adults, which are hard moose to bring down.

The Isle Royale wolves continue to reside in three packs. “The territorial skirmishes seem to have settled,” Peterson said. “There was one lone wolf killed by a pack.”  Peterson also said that all three packs should have litters of pups in late April.

The wolf-moose study is supported by Isle Royale National Park, the National Science Foundation, the Earthwatch Institute, and a number of individual donors.

3/6/03


2002 Summary on the Annual Moose-Wolf Study

HOUGHTON MI -- The lack of winter weather explains many of the changes in the annual wolf-moose survey at Isle Royale National Park, according to Michigan Technological University's Rolf Peterson. Peterson, a professor of wildlife ecology at Michigan Tech, said the 2002 survey counted 17 wolves on the island, as opposed to 19 last year. The island's moose population increased from about 900 last year to 1,100 in 2002.

"The significant factor was a lack of winter," Peterson said, referring to the very light snowfall this year. "Moose were in places where we don't normally see them in the winter--on hillsides and out of the conifer swamps." The Isle Royale wolf-moose survey is the longest running predator-prey study in the world, now in its 44th year. Peterson has conducted the study for the last 32 years. As an island in the middle of Lake Superior, Isle Royale presents a unique opportunity for such research.

Peterson said the wolves suffered a mortality rate of almost 50 percent this winter and that last year's seven pups kept the population near steady. The Isle Royale wolves have formed three packs: the east pack with six members, the Chippewa Harbor pack with five, and the middle pack with four. There are also two single wolves unattached to a pack. Researchers confirmed a confrontation between the east pack and the Chippewa Harbor pack, with the east pack's alpha male killed. Peterson's team found that particular wolf, one of four on the island wearing a radio collar; just offshore in Lake Superior.

"The Chippewa Harbor pack has been pushing the east pack aside and enlarging their territory," Peterson said. "A mild winter is always tough on the wolves and this probably contributed to the confrontation. "We watched about 15 encounters between wolves and moose this winter," he said. "The moose were almost always intimidating, so the wolves didn't bother to attack."

Typically the wolves prey on old moose and young calves. But with little snow cover, the moose could move more freely and avoid the wolves. "Last year, two-thirds of the kills were moose calves," Peterson said. "This year, it was about 20 percent. The wolves had to work hard to find a moose to kill." Peterson also said that all three packs had breeding activity, so he expects three litters of pups to be born in late April.

While the moose have fared relatively well this winter, Peterson says they could begin to have problems this spring and summer. "The trend of warmer temperatures, and a warm, dry spring, could mean an increase in tick infestation of moose," he said. High winds in December also caused a large blow-down of the balsam firs on the western half of the island. The moose prefer these for food. "We lost 16 percent of the fir trees on the western half this winter," he said. "In the last 13 years, about three-quarters of these trees have disappeared. Younger trees can't grow because the moose are eating them. While I expect the moose to increase over the summer, there are some big uncertainties ahead."

The wolf-moose study is supported by Isle Royale National Park, the National Science Foundation, the Earthwatch Institute, and a number of individual donors.

3/11/02


2001 Annual Summary Report

Isle Royale wolves down in numbers -- but eating well

"This winter wolves have the advantage.... This is about as good as it gets for a wolf.'' Dr. Rolf Peterson, Michigan Tech scientist who directs the surveys for the National Park Service. The number of wolves on Isle Royale dropped by one-third over the past year, but wildlife researchers say it's not cause for alarm.

The 31st annual winter survey of wolves on the Lake Superior island revealed 19 wolves, down from 29 one year ago. But those 19 wolves are feasting on an abundance of moose calves, and the wolf packs are enjoying "the best of times'' said Dr. Rolf Peterson, the Michigan Tech scientist who directs the surveys for the National Park Service. Moose now number about 900, up slightly from 850 last year and still rebounding after bottoming out at 500 after the deep snow winter of 1995-96. There is an especially strong crop of calves from 2000, Peterson said.

The decline in wolves can be attributed to inter-pack warfare and a smaller crop of moose calves in 1999. There were fewer calves that year probably because of the hot, dry summer of 1998, Peterson said. It's yet another example of the complex predator/prey relationship on the island that has become a wild laboratory for Peterson and other scientists. The relationship is considered unique because wolves are the only predators and moose the only major prey. Peterson has tracked both populations to see what impact changes in one species have on the other, all in a controlled environment with little human interference. The animals can't leave the isolated island and there are no vehicles, poachers or hunters to affect the population.

During the easy winter of 2000, wolves on Isle Royale weren't able to kill as many moose as they needed. But that problem actually was set in motion 18 months earlier, when weather began affecting moose. The warm summer of 1998 made life difficult for moose because they don't perspire. Instead they become overheated, hyperventilate and have to use a lot of energy to try to keep cool. Those same conditions led to a greater tick infestation the following winter, which further weakened the moose. The result was that fewer than 100 calves were born in 1999 -- important because calves provide the most reliable prey for wolves. With light snow, as in the winter of 1999-2000, moose can move around more easily and are harder to catch, so wolves have 0hard times. This year's heavy snow cover means Isle Royale's wolves are finding easy pickings.

"The 2000 calf crop produced between 200 to 300 animals,'' said Peterson. "And heavy snow makes traveling tough for calves, so this winter wolves have the advantage.... This is about as good as it gets for a wolf.'' The snow in northern Minnesota and Wisconsin's wolf range isn't quite as deep as on the island, but those wolves are probably thriving this winter as well, said Bill Berg, wildlife biologist for the Minnesota Department of Natural Resources in Grand Rapids. Wolves have the same deep snow advantage over deer, their primary food across most of the Northland, as they do on young moose. "In years of very deep snow, the advantage really shifts to the wolf. But I'm not sure we're there yet, not quite,'' Berg said. "I've seen some fawns really floundering, having a hard time getting through the snow, so I'm sure we'll see some fawn mortality. But for most deer, this should be an average winter. Same for most wolves.''

On Isle Royale, if there was an equilibrium between the species, it would be about 30 wolves and 1,000 moose, Peterson said. But, despite the controlled environment, that level is almost never reached -- one of the two species is almost always out of whack. Peterson said the island's West Pack of wolves has been eliminated and its territory taken over by the Middle Pack, which numbers six animals. The six-member East Pack has maintained its territory on the moose-rich east end. "And there is a third pack of just a reproducing pair and one surviving pup that has carved out a territory for itself on the northeast side of the park, and the other packs have respected its territory,'' Peterson said. "There is also another mated pair and two single wolves wandering about the island and making do as best they can.''


2000 Annual Summary Report

Last year, the world’s longest running wildlife research project – the 42nd year of the wolf and moose monitoring program – continued during January and February of 2000, as usual. This annual "Winter Study" project provides the best opportunities for aerial surveying of the wolf and moose populations, with leaves off the trees and snow on the ground.

In 2000, the ISLE ROYALE WOLVES were also doing well. Last winter, the prediction was that the wolves should continue to hold their own in the foreseeable future, according to the completed survey of the island's wolf and moose populations. Park Superintendent Douglas Barnard said the island's wolf packs added five pups to the total population during 1999, but one adult wolf was killed in 2000, leaving the park with 29 wolves last winter. Just two years before, 1998, the population had plummeted to 14, and biologists and park officials were concerned about the species' continued survival. But those concerns seem to have been allayed by the wolves' remarkable ability to withstand nature's challenges.

"In general the island's wolves appear to be in good health, and the packs are experiencing normal reproductive success," said Barnard last winter. "We see no reason why this trend shouldn't continue over the next few years." MTU Wildlife Biologist Dr. Rolf Peterson, who has conducted the annual wolf/moose survey for the National Park Service for the better part of three decades, reported last year that there had been a realignment of pack territories on the island. "Whereas we used to have three packs dividing the island, now there are essentially only two," said Peterson. "The Middle Pack, which had three surviving pups from last year and now numbers 12 animals, has virtually taken over the former West Pack territory. The West Pack's numbers have steadily declined, and they are simply no longer able to defend their territory. There may be a couple of them still alive, but we haven't been able to determine that for certain." He said there are also two other adult pairs roaming the island, as well as a single adult wolf. The East Pack successfully raised two pups from last year and now has 10 members, which continue to control the east end of the island.

Biologists picked up the carcass of one wolf that succumbed to interpack warfare and witnessed the near demise of another. "An independent adult pair of wolves had been roaming the border area between the East and Middle Pack territories for some time," said Peterson. "One day we were flying over that part of the island tracking the signal from a radio collar of a male wolf that indicated the animal had died. When we located the site from which the signal was coming, we saw the Middle Pack standing there. Later we returned to the area and found the carcass of a wolf where the pack had been earlier. It turned out to be the male of the pair that had been using the border area. It had been killed by the Middle Pack.

"Two days later we were flying along the shoreline of the island. We saw a female standing on a rock out in Lake Superior facing 10 wolves from the Middle Pack, who were pacing the shore opposite her. Some of them crossed on rocks to where she was and attacked her several times. We watched this go on for about 45 minutes before she jumped into the water and swam away. She swam about 100 yards up the lake, then came ashore. As soon as she was on land, the wolves from the Middle Pack attacked her in force and appeared to have killed her. After a while they left her body and moved off into the woods, and we returned to camp to refuel our plane and get a pack to haul out her carcass.

"When we returned to the site nearly three hours after the attack, we noticed a lone male from the Middle Pack casually walking along the shoreline trail. He had not been with the pack earlier and apparently was trying to catch up with them. When he came to the site of the battle, he became very uneasy and then noticed the body of the female. To our surprise, when he approached her, she raised her head and looked at him. After a while she was able to pull her forelegs up under her, but she was still unable to stand. The male wolf then went over to a nearby rock where he lay down and watched the female, who was in heat.

"We had planned on landing at a nearby inland lake and then hiking to the shore to pick up the female's body. When we saw she was alive, we left the area and went to check on other wolves. When we returned two hours later to check on the situation, both animals were gone. We saw them again five days later about a half-mile from the battle scene. They were standing atop a little knob, and we were able to see clearly that the male was licking some wounds on the female's neck. We never saw them again, so we don't know for sure whether or not she made it."

Peterson also said last year that the Middle Pack was controlling about three-fourths of the park's area, and by taking over the west end of the island they have increased the number of moose available to them for food. The East Pack, however, while having a smaller territory, still maintained control over that part of the island with the greatest number of moose.

Isle Royale's moose population has increased slightly from '99 to '00, from 750 to 850 in 2000, according to Peterson. He said then that the population seemed stable, but calves are scarce so overall growth potential of the herd appears minimal. "I was surprised to see so few calves," he said. "It's one of the lowest levels we've ever recorded. We don't know why, but it could be weather-related." He said hot, dry summers such as Isle Royale experienced in 1998 create stress for moose. "Moose don't perspire," he explained, "so hot weather causes them to overheat, and they don't eat as well as they should and as a result, aren't as healthy. Studies in Norway have shown that moose killed by hunters after a hot summer have greatly reduced body weights compared to those killed following a cool, moist summer which moose prefer. He said early spring and hot summers also result in higher tick production and greater infestation on moose.

"[1999] was almost a non-winter on Isle Royale," said Peterson. "So for the moose's sake, we should hope for a cool, moist summer. The short-term success of the island's herd probably depends as much on the weather as anything else."

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